Nanney, David Ledbetter, 1925-

The experimental ciliatologist David L. Nanney spent much of his career studying the protozoan Tetrahymena. Under Tracy M. Sonneborn at Indiana University, he completed a dissertation in 1951 on the mating habits of Paramecium, but soon after joining the faculty at the University of Michigan, he turned his attention to Tetrahymena. During his subsequent career in Ann Arbor (1951-1959) and at the University of Illinois (1959-1991), Nanney made a series of fundamental contributions to the cytology, genetics, developmental biology, and evolution of ciliates, influencing the work of other biologists such as Joe Frankel, Janina Kaczanowska, Linda Hufnagel, and Nicola Ricci. Since his retirement in 1991, Nanney has remained in Urbana.

The Nanney Papers include a dense run of professional correspondence with ciliatologists, geneticists, students and colleagues regarding his pioneering research on ciliates and other professional matters. Of particular note is an extensive correspondence with Sonneborn, accompanied by several biographical essays written after Sonneborn’s death, and a large body of correspondence of the controversial reorganization of the biological sciences departments at the University of Illinois in the 1970s. The collection also includes a selection of Nanney’s writings and a handful of photographs.

Background on David L. Nanney

The ciliatologist David Ledbetter Nanney was born in Abingdon, Va., on October 10, 1925, the son of T. Grady and Pearl Ledbetter Nanney. As an infant, David’s father, a Baptist minister, moved from the Blue Ridge Mountains to the oil boom town of Wewoka, Oklahoma, capitol of the Seminole Nation. Although not highly educated in a formal sense, his family was nevertheless strongly oriented toward learning. After graduating high school and being excluded from military service due to the effects of childhood ailments, Nanney entered Oklahoma Baptist University in Shawnee, Okla., with the class of 1946. Studying literature and philosophy, he racked up a mixed academic record, and when his hopes of securing funding to continue his studies in the humanities fell by the wayside, he seized the opportunity to enter a field where funding was deeper and the need for graduate students more acute: zoology.

Following this rather uncommon path into science, Nanney entered Indiana University in the fall of 1946. In the post-war years, Indiana was home to a vibrant, often fractious collection of biologists, including the future Nobel laureates Herman J. Muller and Salvador Luria and the distinguished geneticist, Tracy M. Sonneborn. Despite Nanney’s meager background in biology, Sonneborn agreed to become his research advisor, and fairly soon thereafter, Nanney became immersed in research on Paramecium aurelia, the ciliate with which Sonneborn was most closely identified. The Sonneborn lab included an exceptionally talented group of researchers, including John Preer (Sonneborn’s first graduate student), Richard Siegel, P.K. Chao, and Ruth Dippell, all of whom went on to notable careers, and the group of experimental ciliatologists grew to include Ko Hiwatashi, Janine Beisson, Renzo Nobili, and Klauss Heckmann, notable figures in the study of Paramecium in Japan, France, Italy, and Germany, respectively. James Watson, a Luria student and future Nobel laureate, was another frequent associate.

During Nanney’s years a grad student, Indiana was at the center of vigorous debate over the nature of the evolutionary process, with Sonneborn’s theory of cytoplasmic inheritance pitted against Muller’s “nucleic hegemony.” Nanney’s contributions to this debate were somewhat unexpected. In his dissertation research on quantitative differences in kappa particles between two mating types in Paramecium, he concluded that some form of epigenetic control system (to use a term that would not be coined until several years later), rather than cytoplasmic inheritance, was at work in regulating activity states in nucleic information. Reluctantly, and graciously, Sonneborn agreed. The resulting paper on epigenetic control systems, one of the first significant contributions of his career, earned Nanney a place in Proceedings of the National Academy of Science. Sonneborn, without doubt, was the most significant influence on Nanney’s career, not only serving as his entry point into the field, but as a mentor, sounding board, and model on whom to build a research career.

Shortly after receiving his doctorate in June 1951, Nanney married Jean Kelly, a graduate student in piano and reviewed his career options. Though tempted by the offer of a National Research Council Postdoctoral Fellowship to work with Max Delbruck at CalTech and delve into phage genetics, Nanney declined in favor of a permanent position in the Department of Zoology at the University of Michigan, and shortly after his wedding, he packed up and moved.

By 1952, Nanney was searching for a new organism to develop into a model for genetic research, and through a colleague, Del Elliott, he began to work with Tetrahymena, the organism with he remained most closely associated throughout the rest of his career. His early research centered on the isolation of mating types, cytological studies of conjugation, and the genetics of mating type differentiation, laying the sort of solid foundation that has made Tetrahymena a model. He did, however, feel somewhat isolated in Michigan, sharing little with his colleagues, with Sonneborn continuing to serve as his primary scientific ally and critic. At Michigan, Nanney also became interested in reforming undergraduate biological education to keep pace with rapid developments in the field. His ideas for curricular reform at Michigan, however, were consistently thwarted by what he saw as an ossified system. In this aspect, too, Michigan seemed less than an ideal match.

When his first sabbatical cycle rolled around in 1958, Nanney was forced to turn down a Fulbright Fellowship to work with Boris Ephrussi in Paris (fearing that it would make him eligible for the draft), but he returned to thoughts of CalTech, and received funds to spend a year working there with Ray Owen. Removed from Ann Arbor, Nanney’s frustrations with the pace of curricular reform reached a head, and when offered a position at the University of Illinois, where the varied departments that dealt with biology had recently been consolidated into a new School of Life Sciences (SOLS), he jumped. The presence of Luria, Marcus Rhoads, and Sol Spiegelman sweetened the deal.

Nanney’s arrival at Illinois coincided with the major upsurge of support for science education in the wake of Sputnik and the Cold War, giving him an opportunity to act upon his curricular ideas. The National Science Foundation provided significant funding for Illinois to re-emphasize the undergraduate laboratory experience, stressing a model curriculum built around the “mantra” of the Cell, the Organism, and the Population. While the Illinois project never became the model for others to follow as it was intended, the new curriculum was adopted by the honors program in SOLS and was considered a long-term success. In addition to his curricular work at Illinois, Nanney also served as a member of the Biological Sciences Curriculum Study group, which resulted in the publication of his widely-adopted textbook The Biology of Cells.

With the rapid evolution of the life sciences since the 1960s, reorganizations of the schools and departments became increasingly necessary, and increasingly common. Nanney was involved in several later initiatives to reformulate biological instruction at the university and reorganize the school, taking part in a systematic study of how SOLS faculty related to one another in theory and practice. The result was a series of departmental realignments and reorganizations. Returning from a Humboldt Fellowship in Munster, Germany, in 1984, Nanney discovered that his department, then called Genetics and Development, had actually been disbanded. Rather than return to the reorganizational fray, he accepted an opportunity to join the interdisciplinary Center for Advanced Studies, which eventually spawned a Science and Technology Studies program in which Nanney was active. Nanney has subsequently written at length about the history of protistology, Sonneborn, ciliate genetics, and evolutionary biology.

In Illinois, Nanney built an active research program in Tetrahymena. From the analysis of cortical pattern during the 1960s and 1970s, he and his lab studied genic control, cortical variation and the constancy that “lurks behind this variation,” and by the 1970s, he and his students Dennis Borden and Elizabeth Miller, along a colleague Greg Whitt, began work on isozymes to sort out species in Tetrahymena. The precision he was able to bring in distinguishing strains paved the way for valuable work in the study of molecular evolution, including his later work comparing RNA sequences. Nanney’s publications contributed to a growing debate over the rates of phenotypic versus molecular evolution, with Nanney arguing that in ciliates, at least, evolution at the molecular and morphological levels are relatively uncoupled.

During his career, Nanney served on editorial boards of several journals, including the Journal of Experimental Zoology, Developmental Genetics, Journal of Protozoology, American Naturalist, and Evolution, and he was active in a number of professional organizations, including the American Genetic Association (president, 1982), Genetics Society of America, Society of Protozoologists, and the International Society for Evolutionary Protistology. Since his retirement in 1991, he has remained in Urbana and continues to write on the history of his discipline.

Contents of Collection

Series Descriptions

Series 1. Correspondence	1950-2008	10 boxes
Beginning in the late stages of his doctoral research, the correspondence in Series 1 offers a remarkably dense, sustained perspective on the research life of a significant protistologist. After Sonneborn’s death in 1981, Nanney was called upon both to write and collect memoirs to commemorate the life and scientific contributions of one of the great geneticists of the mid-century. In addition to the extensive and valuable correspondence between Nanney and Sonneborn, the series includes correspondence with a number of other geneticists and protistologists who contributed to a commemorative volume on Sonneborn. Furthemore, the collection includes correspondence with Sonnerborn’s son David, his wife Ruth, and daughter Joan Smith-Sonneborn.
Series 2. Teaching and Writing	1946-2008	3 boxes
Although the series does not by any means include a complete run of Nanney’s publications, it does include several of his key publications, a relatively good representation of his writing on the history of science, drafts of a handful of lectures, and records of grants that help flesh out the evolution of his career and his thoughts on his discipline. The relatively large quantity of material pertaining to the University of Illinois includes some routine paperwork, but consists largely of an interesting set of correspondence, notes, and memos relating to several administrative efforts in the 1970s and 1980s to reorganize the biological sciences departments at the university to reflect larger shifts in the discipline, and materials relating to parallel efforts to improve undergraduate education, particularly writing, in biology.

Inventory of Collection

Series 1. Correspondence	1950-2008	10 boxes
Abraham, Irene	1982		Box 1
Adoutte, Andre	1978-1988		Box 1
Alexander von Humboldt Stiftung	1983-1985		Box 1
Alcorn, Robert D.	1966		Box 1
Alfert, Max	1963		Box 1
Allen, John M.	1966		Box 1
Allen, Robert D.	1969		Box 1
Allen, Sally	1958-1959		Box 1
Allen, Sally	1960		Box 1
Allen, Sally	1961		Box 1
Allen, Sally	1962		Box 1
Allen, Sally	1963		Box 1
Allen, Sally	1964-1965		Box 1
Allen, Sally	1970-1981		Box 1
Allen, Sally and Eduardo Orias, Introduction: a tribute to David L. Nanney, an experimental ciliatologist. Developmental Genetics 13	1992		Box 1
Alliance for Undergraduate Education	1988		Box 1
Alpert, Daniel	1969-1988		Box 1
Alexander, Gordon	1958		Box 1
Ambrose, Harrison	1973-1978		Box 1
American Academy of Arts and Sciences	1978-1985		Box 1
American Association for Cancer Research	1988-1989		Box 1
American Genetic Association	1983-1990		Box 1
American Genetic Association. Frank N. Meyer Medal	1981		Box 1
Frederick G. Meyer
American Genetic Association. Frank N. Meyer Medal	1982		Box 1
Desmond D. Dolan
American Institute of Biological Sciences	1965-1970		Box 1
Ammerman, Dieter	1964-1984		Box 1
Angeloff, L.	1962		Box 1
Bacci, G.	1962		Box 1
Barth, Friedrich G.	1984		Box 1
Batson, B. S.	1983-1985		Box 1
Beale, Geoffrey H.	1960-1969		Box 1
Beam, Carl A.	1986-1991		Box 1
Behnke, Frances L.	1980-1981		Box 1
Behnke, John A.	1980-1983		Box 1
Beisson, Janine	1964-1978		Box 1
Berenbaum, May	1999		Box 1
Berger, Jacques	1975-1985		Box 1
Berger, James D.	1974-1978		Box 1
Berrrill, N. John	1970-1972		Box 1
Blackburn, Elizabeth	1981-1983		Box 1
Bleyman, Lea	1963-1975		Box 1
Boell, E. J.	1983		Box 2
Bogart, Elizabeth	1962		Box 2
Borden, Dennis	1973-1974		Box 2
Borst, Piet	1974-1982		Box 2
Boutin, C. F.	1992		Box 2
Bovee, Eugene C.	1983		Box 2
Bradbury, Elizabeth	1984-1988		Box 2
Brink, R. Alexander	1961		Box 2
Brothers, Janice	1979		Box 2
Brown, Susan	1998		Box 2
Bruns, Peter	1971-1982		Box 2
Brygoo, Yves	1980-1981		Box 2
Burian, Richard M.	1985-1986		Box 2
Butzel, Henry M.	1982		Box 2
Bychkovskaya, Irina Borisovna	1983		Box 2
Byers, Thomas J.	1981		Box 2
Byrne, Barbara	1975		Box 2
Campbell, Gregory R.	1976		Box 2
Carlson, Elof Axel	1987		Box 2
Carlson, Peter F.	1971		Box 2
Carson, Hampton L.	1967		Box 2
Caspari, Ernst W.	1972		Box 2
Cech, Thomas	1979-1981		Box 2
Cesario, Albert	1981		Box 2
Chao, Pao Kuo	1951-1987		Box 2
Chen, Sheng Sheng	1977-1979		Box 2
Chen, T. T.	1950-1966		Box 2
Chen, Yueh Tseung	1981		Box 2
Cho, Paula	1972		Box 2
Clark, Gordon M.	1956-1957		Box 2
Clark, William A.	1965		Box 2
Cleffmann, Gunther	1983-1989		Box 2
Cleland, Ralph E.	1967-1974		Box 2
Cohen, Larry W.	1962-1976		Box 2
Cold Spring Harbor Laboratory	1983-1984		Box 2
Coleman, Annette W.	Undated		Box 2
Collins, Robert L.	1976		Box 2
Conference on Extrachromosomal Heredity	1958		Box 2
Confrey, Eugene A.	1965		Box 2
Corliss, John O.	1952-1967		Box 2
Corliss, John O.	1973-1979		Box 2
Corliss, John O.	1980-1990		Box 2
Corliss, John O., Selected references on protozoa	1963		Box 2
Creager, Angela N. H.	1988		Box 2
Cronkite, Donald L.	1978		Box 2
Crow, James F.	1969-1987		Box 2
Cutler, Preston	1963		Box 2
Dalby, Arthur	1958		Box 2
Davidson, John A.	1978		Box 2
Davis, Elizabeth	1982		Box 2
Dawson, Peter S.	1969		Box 2
Dawson, William R.	1973		Box 2
Denton, Michael	2001		Box 2
De Terra, Noel	1970		Box 2
Developmental Genetics (Journal)	1978-1988		Box 2
Din, Nanni	1977		Box 2
Dini, Fernando	1984-2001		Box 2
Dippell, Ruth V.	1956-1982		Box 2
Dippell, Ruth V., Memorial resolution… Tracy M. Sonneborn for inclusion in the Faculty Council minutes	18981		Box 2
Doerder, F. Paul	1968-1977		Box 3
Doerder, F. Paul	1978-1985		Box 3
Drill, Victor	1970		Box 3
Dryl, Stanislaw	1972-1989		Box 3
Duina, Andrea	1991-1992		Box 3
Eck, Michael John	1986		Box 3
Ehret, Charles	1964		Box 3
Elliott, A. M.	1974		Box 3
Ellstrand, Norman	1974-2000		Box 3
Engberg, Jan	1983		Box 3
Ephrussi, Boris	1955-1962		Box 3
Erdstrom, Jan Erik	1969		Box 3
Esposito, Fulvio	1977-1984		Box 3
Faberge, A. C.	1986		Box 3
Fava, George	1973		Box 3
Ferguson, Beth	1979		Box 3
Fine, Sidney	1971		Box 3
Finger, Irving	1969-1971		Box 3
Fitch, Walter M.	1988		Box 3
Fleck, Earl William	1976		Box 3
Flecker, Emmanuel A.	Undated		Box 3
Flint, Franklin F.	1969		Box 3
Forrest, Hugh S.	1978		Box 3
Franceschi, Tina	1963-1992		Box 3
Frankel, Arthur I.	1969		Box 3
Frankel, Joseph	1963-1972		Box 3
Frankel, Joseph	1973-1974		Box 3
Frankel, Joseph	1975-1979		Box 3
Frankel, Joseph	1980-1984		Box 3
Frankel, Joseph	1985		Box 3
Frankel, Joseph	1986-2008		Box 3
Freeman, Gary	1988		Box 3
Freiburg, Manfred	1986		Box 3
Futrelle, Robert P.	1985		Box 3
Gall, Joseph	1976-1985		Box 3
Gates, Michael A.	1975		Box 3
Genermont, Jean	1979		Box 3
Genetics (Journal)	1980		Box 3
Genetics Society of America	1967-1985		Box 3
Genetics Society of America Conference (Urbana, Ill. : 1986)	1985		Box 3
Genetics Student	1977		Box 3
George, Brother L.	1964		Box 4
Getz, Lowell	1965-2005		Box 4
Gibson, Ian	1965		Box 4
Gio-Ardez, Raul	1977-1997		Box 4
Gladiszewski, Maria	1981-1986		Box 4
Glass, Bentley	1987		Box 4
Goldstein, Wayne M.	1973		Box 4
Goldwasser, Edwin L.	1980-1991		Box 4
Good, Phillip I.	1976		Box 4
Goodwin, Carol	1963		Box 4
Gorovsky, Martin	1971-1985		Box 4
Gortz, Hans Dieter	1985		Box 4
Grant, Carroll W.	1958		Box 4
Grass, Frank	1971-1976		Box 4
Grassmick, Robert A.	1979		Box 4
Graves, Lynn	1972		Box 4
Greenburg, A. H.	1981		Box 4
Grimes, Gary W.	1979		Box 4
Grobstein, Clifford	1961-1966		Box 4
Grula, E. A.	1976-1977		Box 4
Gunslaus, I. C.	1967		Box 4
Haller, Gerard de	1984		Box 4
Halvorson, Halvor	1962		Box 4
Hammersmith, Robert	1982		Box 4
Hartman, Hyman	1983		Box 4
Harwood, J. H.	1987		Box 4
Hayes, D. H.	1977-1979		Box 4
Heckmann, Klaus	1969-1982		Box 4
Heckmann, Klaus	1983-1999		Box 4
Henderson, J. Donald	1965		Box 4
Henshaw, Robert E.	1969		Box 4
Higashinakagawa, Toru	1980		Box 4
Hildebrand, Milton	1964		Box 4
Hirsch, Jerry	1967		Box 4
Hiwatashi, Koichi	1966-1983		Box 4
Hogg, James F.	1979		Box 4
Holmquist, Gerald P.	1983-1984		Box 4
Holz, Geoge G.	1958		Box 4
Hutner, Seymour H.	1977-1985		Box 4
Inoki, Shozo	1954-1955		Box 4
International Congress of Cell Biology XI	1963-1964		Box 4
International Congress of Protozoology, 6th (Sonneborn Symposium)	1981		Box 4
International Society for Evolutionary Protistology	1991		Box 4
John Wiley and Sons	1960-1961		Box 4
John Wiley and Sons	1962-1967		Box 4
Johmann, Carol A.	1974		Box 4
Jones, Garth W	1977		Box 4
Juckett, Robert S.	1979		Box 4
Juergensmeyer, Elizabeth B.	1969-1981		Box 4
Kaczanowska, Janina	1972-1994		Box 4
Kaczanowski, Andrzej	1971-1974		Box 4
Kaczanowski, Andrzej	1975-1979		Box 4
Kaczanowski, Andrzej	1980-1984		Box 4
Kaczanowski, Andrzej	1985-1989		Box 4
Kaczanowski, Andrzej	1990-2000		Box 4
Kallio, Reino	1973		Box 5
Kaluzynski, Stanislaw	1984		Box 5
Kaneda, Masao	1963		Box 5
Kaney, Anthony R.	1982		Box 5
Kansas State University	1965-1970		Box 5
Katashima, Ryo	ca.1965		Box 5
Kazubski, Stanislaw	1982		Box 5
Keller, Evelyn Fox	1997		Box 5
Kelsus, A. S.	1971		Box 5
Kendeigh, Charles	1973		Box 5
Kevles, Daniel J.	1986-1987		Box 5
Kimball, Richard F.	1953-1963		Box 5
King, Charles E.	1982		Box 5
Kinnamon, Keneth	1975		Box 5
Kitzmiller, James B.	1958-1976		Box 5
Koffler, Henry	1971		Box 5
Koizumi, Sadaaki	1962		Box 5
Krawiec, Steven	1976		Box 5
Kresin, Daniel	1978-1993		Box 5
Kucherlpati, Raju	1972-1975		Box 5
Kuhn, Barbara C.	1982-1983		Box 5
American Genetic Association
Kung, Ching	1975-1979		Box 5
Lanier, Lyle	1965		Box 5
Lanners, Norbert	1972		Box 5
Larsen, Joseph R.	1972-1981		Box 5
Lederberg, Joshua	1978-1981		Box 5
Tracy M. Sonnerborn
Lefevre, George	1980		Box 5
Leick, Vogn	1979		Box 5
Leiner, Michael	1963		Box 5
Leone, Charles A.	1962		Box 5
Levine, Norman	1972		Box 5
Lewis, Herman W.	1972		Box 5
Lewontin, Richard	1961-1966		Box 5
Lieb, Margaret	1972		Box 5
Lindholm, Tore	1980		Box 5
Includes photograph
Lindquist, David L.	1969		Box 5
Little, James W.	Undated		Box 5
Loefer, John B.	1961		Box 5
Lom, Jiri	1973-1974		Box 5
Lopez-Ochoterena, Eucario	1980		Box 5
Luce, William M.	1959		Box 5
Luporini, Pierangelo	1977-1988		Box 5
Luria, Salvador E.	1959		Box 5
Lynn, Denis H.	1971-2003		Box 5
Lyo, John	1994		Box 5
Maly, Roland	1961		Box 5
Maresca, Bruno	1986		Box 5
Margolin, Paul	1956-1958		Box 5
Margulis, Lynn	1971		Box 5
Markert, Clement L.	1962-1985		Box 5
Martindale, Duane	1989		Box 5
Matsushiro, Aizo	1957-1958		Box 5
McDonald, Barbara B.	1957-1965		Box 5
McKeachie, Wilbert J.	1954-1989		Box 5
McKoy, J. Wynne	1968-1969		Box 5
McKoy, J. Wynne	1970-1973		Box 5
McKoy, J. Wynne	1976-1979		Box 5
McKoy, J. Wynne, Recombination in Tetrahymena thermophila (Dissertation, Univ. of Illinois)	1976		Box 5
McQuade, A. B.	1989		Box 6
Medvedev, Zh. A.	1963		Box 6
Meins, Frederick	1979-1983		Box 6
Melton, Douglas	1974		Box 6
Meredith, A.	1980-1981		Box 6
Miceli, Cristina	1985-1992		Box 6
Mikami, Kazuyuki	1981		Box 6
Milkman, Roger	1957-1985		Box 6
Miller, Arnold R.	1980		Box 6
Miller, James G.	1971		Box 6
Miyake, Akio	1976-1985		Box 6
Mobley, Dale O.	1994-2001		Box 6
Morin, Peter	2006-2008		Box 6
Muhlbock, O.	1962		Box 6
Murphy, Donald G.	1970		Box 6
Myohara, Koji	1979		Box 6
Myohara, Maroko	1981		Box 6
Najamuzzaman	1969		Box 6
Nanney laboratory	1983-1984		Box 6
Correspondence with laboratory during sabbatical at the University of Munster
National Research Council	1965-1989		Box 6
Neel, James V.	1982		Box 6
Nelsen, E. Marlo	1978		Box 6
New York Academy of Sciences	1969		Box 6
Ng, Stephen	1976-1985		Box 6
Nielsen, Henrik	1983-1986		Box 6
Nobili, Renzo	1979-1994		Box 6
North Texas State University	1981		Box 6
Nyberg, Dennis	1972-1973		Box 6
Nyberg, Dennis	1974-1978		Box 6
Nyberg, Dennis	21979-2001		Box 6
Ohio State University	1969-1973		Box 6
Oklahoma Baptist University	1972-1996		Box 6
Oregon State University	1967		Box 6
Orel, V.	1987		Box 6
Orias, Eduardo	1958-1970		Box 6
Orias, Eduardo	1973-1979		Box 6
Orias, Eduardo	1980-1985		Box 6
Osmundsen, John A.	1962		Box 6
Owen, Ray D.	1954-1995		Box 6
Paigen, Kenneth	1963-2005		Box 6
Palmer, Graham A.	1981		Box 6
Palmer, R. R.	1965		Box 6
Pang, Yan Bin	1981-1986		Box 6
Paramecium Conference, 2nd (Bloomington, Ind.)	1951		Box 6
Pardi, C. L.	1986		Box 6
Patterson, David J.	1964-1984		Box 6
Payne, Fernandus	1975		Box 6
Perkins, David D.	1965-1980		Box 6
Perlman, Bruce S.	1972		Box 6
Phelps, Austin	1954-1956		Box 7
Phi Kappa Phi	1967		Box 7
Phillips, Ruth	1969-1972		Box 7
Pierce, James M.	1973		Box 7
Portnoy, Stephen	1980		Box 7
Preer, John R.	1952-1995		Box 7
Preer, John R., Nomination of Tracy M. Sonneborn for the Nobel Prize	Undated		Box 7
Preer, John R., Tracy Morton Sonneborn, October 19-1905-January 26, 1981	1981		Box 7
Preparata, Rose Marie	1978-1981		Box 7
Prescott, David	1980		Box 7
Quastler, Henry	1951		Box 7
Rao, M. V. Narasinha	1958-1964		Box 7
Rasmussen, Leif	1984-1992		Box 7
Ray, Charles	1956-1958		Box 7
Reeve, John N.	1979		Box 7
Reissig, Jose L.	1975-1977		Box 7
Remondini, David J.	1980-1986		Box 7
Rhoades, Marcus	1985		Box 7
Ricci, Nicola	1977-1979		Box 7
Ricci, Nicola	1980-1984		Box 7
Ricci, Nicola	1985-2001		Box 7
Ricci, Nicola. In memoriam	2001		Box 7
Rice, Stanley	2001		Box 7
Richmond College, CUNY	1971		Box 7
Riley-Kresin, Catherine E.	1981		Box 7
Ringertz, Nils	1979-1984		Box 7
Roberts, Dennis	Undated		Box 7
Roderick, Tom	1961		Box 7
Roman, Herschel L.	1981		Box 7
Roque, Madeline	1956		Box 7
Rotheim, Minna B.	1981		Box 7
Rubin, Harry	1989		Box 7
Rudzinska, Maria A.	1957		Box 7
Russell, Elizabeth S.	1975		Box 7
St. Marys College	1964		Box 7
Salceda, Victor M.	1981		Box 7
Sand, Seaward A.	1961		Box 7
Sangsoda, Siddhichoke	1976		Box 7
Sapp, Jan	1983-1992		Box 7
Sargent, M. L.	Undated		Box 7
Satir, Birgit	1971-1983		Box 7
Savard, Edward	1972		Box 7
Schewe, Edgar R.	1978		Box 8
Schloegel, Judy Johns	1995-1998		Box 8
Schloegel, Judy Johns, From anomaly to unification: Tracy Sonneborn and the species problem in protozoa, 1954-1957. Journal of the History of Biology 32	1999		Box 8
Schmidt, Helmut	1986-1996		Box 8
Schmitt, Gail K./	2001		Box 8
Schull, William J.	1982		Box 8
Schulmann, Murray	1963		Box 8
Schuster, Frederick L.	1969		Box 8
Schwartz, Viktor	1979		Box 8
Serre, Arlette	1961		Box 8
Seyfert, Hans-Martin	1976-1978		Box 8
Seyfert, Hans-Martin	1979-1988		Box 8
Shi, Xin Bai	1982		Box 8
Shulman, Richard	1969		Box 8
Siegel, Richard	1960-1981		Box 8
Siegel, Richard	1997-2006		Box 8
Siegel, Richard, A personal panegyric (on the life of Tracy M. Sonneborn)	1981		Box 8
Silverman, Paul H.	1965		Box 8
Simon, Ellen M.	1973-1988		Box 8
Simon, Ellen M. et al., The Tetrahymena pyriformis complex of cryptic species	2006		Box 8
Slater, Donal W.	1970-1971		Box 8
Small, Eugene B.	1965		Box 8
Smith, Hobart	1969		Box 8
Smith-Sonneborn, Joan	1980-1983		Box 8
Smithies, Oliver	1975		Box 8
Snell, William J.	1969-2005		Box 8
Society of Protozoologists	1962-1988		Box 8
Sogin, Mitchell	1985		Box 8
Soldo, Anthony T.	1976		Box 8
Sonneborn, David	1972-1983		Box 8
Sonneborn, Ruth	1982		Box 8
Sonneborn, Tracy M.	1951-1952		Box 8
Sonneborn, Tracy M.	1953		Box 8
Sonneborn, Tracy M.	1954		Box 8
Sonneborn, Tracy M.	1955		Box 8
Sonneborn, Tracy M.	1956		Box 8
Sonneborn, Tracy M.	1957		Box 8
Sonneborn, Tracy M.	1958-1959		Box 8
Sonneborn, Tracy M.	1960-1964		Box 8
Sonneborn, Tracy M.	1965-1967		Box 9
Sonneborn, Tracy M.	1968-1970		Box 9
Sonneborn, Tracy M.	1971-1973		Box 9
Sonneborn, Tracy M.	1974		Box 9
Sonneborn, Tracy M.	1975-1977		Box 9
Sonneborn, Tracy M.	1978-1980		Box 9
Sonneborn, Tracy M., Autobiography (2)	Undated		Box 9
Sonneborn, Tracy M. Biographical	1978-1981		Box 9
Sonneborn, Tracy M. Biographical: A la memoire de Tracy M. Sonneborn, by Jean-Michel Labouygues	1981		Box 9
Sonneborn, Tracy M. Biographical: Annual Review of Genetics (correspondence)	1981		Box 9
Sonneborn, Tracy M. Biographical: Biographical Memoirs of Fellows of the Royal Society, by G. H. Beale	ca.1981		Box 9
Sonneborn, Tracy M. Biographical: Developmental Genetics	ca.1981		Box 9
Sonneborn, Tracy M. Biographical notes and drafts	ca.1981		Box 9
Sonneborn, Tracy M. Biorgaphical: Yearbook of the American Philosophical Society, by Bentley Glass	1982		Box 9
Sonneborn, Tracy M., Ethical issues arising from the possible uses of genetic knowledge	1973		Box 9
Sonneborn, Tracy M. , Genetics and man’s vision, Proceedings of the American Philosophical Society 109, 4	1965		Box 9
Sonneborn, Tracy M. Honorary degree presentation, University of Munster, by Klaus Heckmann	1979		Box 9
Sonneborn, Tracy M. , Implications of the new genetics for biology and man, AIS Bulletin 13, 2	1963		Box 9
Sonneborn, Tracy M. , My intellectual history in relation to my contributions to science	1978		Box 9
Sonneborn, Tracy M. Remarks of Lee Sonneborn, Gros Louis, James Ebert, John R. PReer, and David Sonneborn	1981		Box 9
Sonneborn, Tracy M. Sonneborn Bibliography	ca.1980		Box 9
Spiegelman, George B.	1966		Box 9
Stadler, David R.	1973		Box 9
Stahl, Franklin	1983		Box 9
Stebbins, G. Ledyard	1978		Box 9
Steffensen, Dale M.	1982		Box 9
Stern, Curt	1969		Box 9
Stern, Herbert	1966		Box 9
Stocum, David	1979		Box 9
Streisinger, George	1983		Box 9
Strick, James	2001-2002		Box 9
Sturtevant, A. H.	1957		Box 9
Suhr-Jessen, Peter B.	1983-1984		Box 9
Suzuki, David	1981		Box 9
Tan, C. C.	1982		Box 10
Tarr, George E.	1974		Box 10
Tartar, Vance	1969-1975		Box 10
Taylor, Saundra	2001		Box 10
Temple University	1966		Box 10
Tepe, William	1964		Box 10
Tetrahymena News	1972-1974		Box 10
Thompson, Guy A.	1978		Box 10
Tiedtke, Arno	1989		Box 10
Tohutski, Linda	1979		Box 10
Trends in Genetics	1984-1990		Box 10
Uhlenhopp, Elliott L.	1978		Box 10
University of California Davis	1969		Box 10
University of Iowa	1963		Box 10
University of Toronto	1981		Box 10
Uzzell, Thomas	2001-2005		Box 10
Van Bell, Craig	1984-1986		Box 10
Van der Kloot, William G.	1981-1982		Box 10
Villiers, T. A.	1965		Box 10
Viterbo College	1969		Box 10
Viza, Dimitri	1978-1979		Box 10
W. W. Norton	1975		Box 10
Wahls, Wayne	1982-1991		Box 10
Wallace, Bruce	1975		Box 10
Washburn, Jan O.	1991		Box 10
Waterman, Talbot H.	1961		Box 10
Watson, James D.	1968-1992		Box 10
Watson, M. R.	1963		Box 10
Wee, Beth	1985		Box 10
Wells, Carolyn	1961		Box 10
Weindruch, Rick	1973-1982		Box 10
Weir, Morton W.	1978		Box 10
Wellnitz, Bill	1971		Box 10
Whitt, Gregory S.	1969-1986		Box 10
Who’s Who in America	1981		Box 10
Wiggenhorn, Anthony W.	1969		Box 10
Williams, Norman E.	1965		Box 10
Willis, Judy	1969-1984		Box 10
Wilson, Katherine S.	1966-1998		Box 10
Wilson, Rob	1998-2005		Box 10
Woese, Carl	1969-1973		Box 10
Woodard, John	1962		Box 10
Woods Hole Marine Biological Laboratory	1966-1989		Box 10
Yao, Meng Chao	1980-1982		Box 10
Zieg, R. G.	1972		Box 10
Ziegler, Bonnie	Undated		Box 10
Zuckerkandl, Emil	1988		Box 10
Unidentified recipients	1970-1989		Box 10
Series 2. Teaching and Writing	1948-2008	3 boxes
Conferences	1969-1989		Box 11
Nanney, David L., Academic profile	1988		Box 11
Nanney, David L., Apologia vitae [autobiography]	1983		Box 11
Nanney, David L., Bibliography	1976		Box 11
Nanney, David L., Biographical			Box 11
Nanney, David L., Book reviews	1978-1996		Box 11
Nanney, David L., Candide in academe [autobiography]	2000		Box 11
Nanney, David L., Candide in academe meets Tracy agonistes [autobiography]	2004		Box 11
Nanney, David L., The cell, the organism, the population: reflections on a career experienced	1986		Box 11
Nanney, David L., Charging windmills [autobiography]	2005		Box 11
Nanney, David L., Correspondence with publishers	1958-1999		Box 11
Nanney, David L., Cortical patterns in cellular morphogenesis Science 160	1968		Box 11
Nanney, David L., Cytogenes and cytotaxis	1966		Box 11
Woods Hole evening lecture series
Nanney, David L., Developmental Genetics 13, 1-3 [festschrift numbers]	1992		Box 11
Nanney, David L., Disciplinary strategies of genetics: a revisionist history	1986		Box 11
Nanney, David L., Drafts of articles	Undated		Box 11
Nanney, David L., Epigenetic factors affecting mating type expression in certain ciliates	1958		Box 11
“Drafts of presentation on epigenetics near the time of the PNAS paper”
Nanney, David L., Experimental Ciliatology: correspondence and reviews	1980-1981		Box 11
Nanney, David L., Frontier connections [manuscript]	1994		Box 11
Read at University of Pisa, Oct. 4, 1994, in return for an honorary degree (honoris causa). Published with minimal changes in Rev. Soc. Mex. Hist. Nat. 47: 201-216.
Nanney, David L., Frontier connections, Rev. Soc. Mex. Hist. Nat. 47: 201-216	1997		Box 11
Nanney, David L., Genetic studies of Paramecium aurelia. Dissertation, Indiana University	1951		Box 11
Nanney, David L., Grants	1960-1969		Box 11
Nanney, David L., Grants	1971-1978		Box 12
Nanney, David L., Grants	1982-1988		Box 12
Nanney, David L., Grants	1990-1994		Box 12
Nanney, David L., Historical perspectives in Tetrahymena genetics	1982		Box 12
Appendix to an NIH grant application
Nanney, David L., I rapporti con la frontiera [Frontier connections]	1994		Box 12
Lecture on receipt of honorary degree from University of Pisa
Nanney, David L., Innocents abroad	1985		Box 12
“Faculty report on tenure of preistrager sojourn to Europe, hosted by Klaus Heckmann in Munster, Germany, with side trips to Italy and Poland. Plus sabbatical report to Dept. of Genetics and Development”
Nanney, David L. Job offers	1965-1982		Box 12
Nanney, David L., Lectures and lecture notes	1984, Undated		Box 12
Nanney, David L., Ciliate systematics Part I: Varieties, syngens, and cryptic species (incomplete)	Undated		Box 12
Nanney, David L., Ciliate systematics Part II: Approaches to tetrahymenid evolution	Undated		Box 12
Nanney, David L., The domestication of Tetrahymena	Undated		Box 12
Nanney, David L., Evolutionary genetics of the ciliates	Undated		Box 12
Nanney, David L., Organisms are technologies too	Undated		Box 12
Nanney, David L., Why study ciliates (incomplete)	1984		Box 12
Munster lecture
Nanney, David L., Organistic technology	Undated		Box 12
Nanney, David L., Notes: Ciliates, why?	Undated		Box 12
Nanney, David L., Notes: The time of ciliate origins (incomplete)	Undated		Box 12
Nanney, David L., Notes on Tetrahymena	Undated		Box 12
Nanney, David L., Metaphor and mechanism: epigenetic control systems reconsidered	1989		Box 12
Original paper presented at symposium on “The epigenetics of cell transformation and tumor development”
Nanney, David L., Nanney clan reunion	1995		Box 12
“Account of a trip made by David and Jean Nanney to Dolgellan, Wales, in 1980s, but written up later”
Nanney, David L., Night thoughts, the confessions of a manic depressive	1946		Box 12
Nanney, David L., Notes	1960s?		Box 12
Nanney, David L., Nucleo-cytoplasmic interaction during conjugation in Tetrahymena. Biological Bulletin 105	1953		Box 12
Nanney, David L., On nocodazole	Undated		Box 12
Nanney, David L., On winning in science BioScience 32, 3	1982		Box 12
Nanney, David L., Personal	1955-1957		Box 12
Nanney, David L., The radiation of eukaryotic protists: string analysis of 5S ribosomal sequences, with Chaeryung Park and Ellen M. Simon	1994		Box 12
Nanney, David L., Research reports	1981-1986		Box 12
Nanney, David L., Retirement scrapbook 1	1991		Box 12
Nanney, David L., Retirement scrapbook 2	1991		Box 13
Nanney, David L., Reviews of articles and grant applications	1965-1996		Box 13
Nanney, David L., Sabbaticals	1973-1985		Box 13
Nanney, David L., Shifting ditypic site analysis: heuristics for expanding the phyolgenetic range of nucleotide sequences, with Rose Marie Preparata, E. B. Meyer and Ellen M. Simon	1987-1988		Box 13
Nanney, David L., String analysis of eukaryotic evolution [grant application Department of Health and Human Services]	1994		Box 13
Nanney, David L., Tetrahymena genetics at Illinois	Undated		Box 13
Nanney, David L., Tracy agonistes: personal interactions: footnotes to a biography of T. M. Sonneborn	1982		Box 13
Photographs	1955-1957		Box 13
Gordon Research Conference: Molecular Biology of ciliated protozoa, Laconia, N.H.: Group shot of participants	1989		Box 12
ISEP 7 [International Society for Evolutionary Protistology, 7th Conference, London, England]: Group shot of participants	1987		Box 12
Sonneborn, Tracy M., standing in laboratory, photo by Dellenback	ca.1970s	Silver print	Box 12
Sonneborn, Tracy M., seated at desk	ca.1970s	3 black and white prints	Box 12
One copy autographed
Sonneborn, Tracy M. and wife	ca.1980s	Color print	Box 12
Ricci, Nicola, with tulip in his teeth	1980s	Color print	Box 12
University of Illinois: Economic realities and ecological strategies. Presentation to the Priorities Committee	1988		Box 13
University of Illinois: Administrative	1969-1992		Box 13
University of Illinois: Appointment to faculty	1959		Box 13
University of Illinois. Department of Genetics and Development	1974-1988		Box 13
University of Illinois. Faculty Advisory Committee	1987-1988		Box 13
University of Illinois. School of Life Sciences	1972-1988		Box 13
University of Illinois. School of Life Sciences academic history, by David L. Nanney	1982		Box 13
University of Illinois: Undergraduate education	1961-1992		Box 13
University of Illinois: An undergraduate laboratory teaching facility	ca.1962		Box 13